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Resources

• Animalogic — How to find your own tardigrades (video)
• Bennett, S. (2019) — Tardigrades conquered space. Now, they could save the global vaccine and blood supply. NOVA | PBS
• Bertolani et al. (2004) — Experiences with dormancy in tardigrades. Journal of Limnology
• Bino et al. (2024) — Possible roles of CAHS proteins from Tardigrade in osmotic stress tolerance in mammalian cells
• Bittel, J. (2016) — Tardigrade protein helps human DNA withstand radiation. Nature
• Boothby et al. (2017) — Tardigrades use intrinsically disordered proteins to survive desiccation. Molecular Cell
• EarthSky (2021) — Why NASA sent tiny water bears into space
• Front Line Genomics — Everything you need (and want) to know about tardigrades
• Guidetti & Rebecchi (Eds.) (2018) — Water bears: The biology of tardigrades. Springer
• Hashimoto et al. (2016) — Extremotolerant tardigrade genome and improved radiotolerance of human cultured cells by tardigrade-unique protein. Nature Communications
• Hesgrove et al. (2021) — Tardigrade CAHS proteins act as molecular Swiss army knives to mediate desiccation tolerance through multiple mechanisms (preprint)
• Hvidepil & Møbjerg (2023) — New insights into osmobiosis and chemobiosis in tardigrades. Frontiers in Physiology
• Kasianchuk et al. (2023) — The biomedical potential of tardigrade proteins: A review. Biomedicine & Pharmacotherapy
• LaFaro, A. (2019) — The survivors. UNC Research
• Lim et al. (2024) — Tardigrade secretory proteins protect biological structures from desiccation. Communications Biology
• maecenium.org — Cool facts about tardigrades
• Nelson, Guidetti & Rebecchi (2010) — Tardigrada, in Ecology and classification of North American freshwater invertebrates
• Nguyen et al. (2022) — Trehalose and tardigrade CAHS proteins work synergistically to promote desiccation tolerance. Communications Biology
• Packebush et al. (2023) — Natural and engineered mediators of desiccation tolerance stabilize human Blood Clotting Factor VIII in a dry state. Scientific Reports
• Phylum Tardigrada — Lucid Central
• PREreview (2022) — Review of "Tardigrade CAHS proteins act as molecular Swiss army knives..."
• Sanchez-Martinez et al. (2024) — Labile assembly of a tardigrade protein induces biostasis. Protein Science
• Sanchez-Martinez et al. (2023) — The tardigrade protein CAHS D interacts with, but does not retain, water in hydrated and desiccated systems. Scientific Reports
• Sofia, M. (2017) — How one of the world's toughest creatures can bring itself back to life. NPR
• Tanaka et al. (2022) — Stress-dependent cell stiffening by tardigrade tolerance proteins that reversibly form a filamentous network and gel. PLOS Biology
• Tardigrada (Water bears) — Invertebrate Life, BYU-Idaho
• Wikipedia — Tardigrade
• Wolf, Minsky & Fass (2019) — The tardigrade damage suppressor protein binds to nucleosomes and protects DNA from hydroxyl radicals. eLife
• Yagi-Utsumi et al. (2021) — Desiccation-induced fibrous condensation of CAHS protein from an anhydrobiotic tardigrade. Scientific Reports
• Zhang & Zhang (2025) — Tardigrades' cytoplasmic abundant heat soluble proteins serve as membrane protectors during dehydration (preprint)